It's a somewhat difficult time to answer this question, honestly. I'll do my best to give you a reasonable response.
#1 Inglourious Basterds -- This is the easiest to answer. IB was splitting kills with Dark Ascension during the boss camping period of last spring and most of the summer, but the opening of Plane of Sky made it clear that DA could not keep up.
#2 Divinity -- It gets hard right away here. The answer to this would have been DA even a month ago, but since the opening of Sky, Divinity has had the most success with earlier content while IB has been dominating Sky.
#3 Wrathful Inquisition -- This might be prematurely stated, but WI appears to be making significant efforts to advance. They've had some success with bosses and planar drops and, from the outside, appear to be competent raiders.
#4 Darkwind -- To the best of my knowledge, this is the new guild formed by the raiders of Remedy, Europa and a few more. They seem to be fairly casual and maybe too nice and polite of people to compete with the more ruthless guilds.
#5 Transcendence -- Early in the server, this was IB's main competition. At some point, Transcendence leadership made a decision that having real lives was more important than doing everything first on Project 1999. They still raid, but much more casually.
So where does that leave Dark Ascension? They appear to be completely in limbo. A lot of people have left the guild and there are frequent complaints and rumors of unhappiness with Durison's leadership. I haven't seen them put together a raid force for a week or so. Whether they'll continue to decline or are just reorganizing remains to be seen.
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